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The term "oak savanna" refers to a plant community
with scattered "open-grown oaks". surrounded by
grasses and forbs. Other terms sometimes used are "oak
opening" and "oak barren." In contrast to an
oak forest, which has a closed canopy (approaching 100%),
the savanna canopy ranges from 10% to 60%. Although oak savannas
are found in other parts of the United States, in southern
Wisconsin and throughout the upper Midwest they were once
a dominant vegetation type. Originally, about 30% of the vegetation
in southern Wisconsin was oak savanna.
It has been estimated that only about 0.01% of the original
oak savanna of Wisconsin still remains. The remaining fragments
of oak savanna thus constitute a rare and highly endangered
community.
Fortunately, Pleasant Valley Conservancy has around 30-50
acres of oak savannas, and restoration work has created one
of the best examples of the oak savanna type in southern Wisconsin.
How do we recognize an oak savanna? The
key characteristic of the oak savanna is the open-grown oak,
a tree that has developed in the open, away from other trees.
Savanna oaks usually have large lower branches, an indication
that they developed without competition from nearby trees.
The photo below, of a bur oak, is an example.
The presence in the habitat of open-grown oaks is a good
indication of a savanna. Even if the habitat was later heavily
invaded by other trees and exotic brush, the open-grown oaks
may still be present.
Large areas of highly degraded but potentially good oak savannas
still exist in the Midwest, especially on the south-facing
slopes and ridge tops. Examination of old air photos, such
as the 1937 ones taken by the U.S. Soil Conservation Service,
show many highly degraded areas that still have open-grown
trees. The existence of these open-grown oaks at Pleasant
Valley Conservancy was a principal reason why restoration
work was undertaken.
Aesthetics of the Oak Savanna Landscape
Surveys of attitudes and perceptions have shown that the oak savanna landscape rates highly in the public mind. Open forests with large, relatively old trees are considered very attractive. Other pleasing factors of the oak savanna include the high plant diversity, the presence of wild flowers, the large openings surrounded by trees, and the extensive vistas and overlooks. These are characteristic of parks, and the park-like setting of oak savannas is appealing, just as it was to early travelers through the Midwest.
| Attractiveness of the Oak Savanna |
| Large trees |
| Open forest understoreys allowing views deep into the forest |
| Herbaceous vegetation on the forest floor |
| Appearance of easy travel through the forest |
| Large openings surrounded by trees |
| Relatively old forests |
| Vegetation diversity |
| Vistas, overlooks, water bodies, unique geological formations |
| Flowering trees or other flowering vegetation |
| Information from Johnson, Paul S. et al. 2009. The Ecology and Silviculture of Oaks, 2nd edition. CABI International, Cambridge, MA |
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| Classic open-grown bur oak in a restored savanna at Pleasant Valley Conservancy. This unit has been burned annually for the past six years. |
The Importance of Fire Probably the most
common reason why the oak savanna community has disappeared
is because of lack of fire. This is a
fire-controlled vegetation. The oaks themselves are fire-resistant,
whereas weedy trees such as walnut, elm, maple, and ash are
not. In presettlement times, fire was used by Native Americans
to keep the woods open and to control woody shrubs. As late
as the 1950s, farmers were using fire for similar purposes
and to bring about early "green-up" of their pastures.
However, protection from fire became an established practice
of the U.S. Forest Service, as well as the Wisconsin Conservation
Commission (the forerunner of the Department of Natural Resources).
Although fire is a real danger in the coniferous forests of
northern Wisconsin, not so for southern oaks. We know now
that it was wrong to prevent fires in the southern oaks. Oaks
do not suffer from the disastrous crown fires that rage through
the coniferous forest. Except under unusual circumstances,
fire in an oak woods is confined to the ground, to the leaf
litter. Oak leaves are unusually susceptible to fire. When
they burn, weedy woody vegetation such as prickly ash, buckthorn,
and honeysuckle is killed, thus keeping the woods open. An
open woods encourages the growth of grasses and flowering
plants, which are part of the glory of an oak savanna.
Ideas have changed completely about fire in the woods. Without
fire, restoration of an oak savanna is difficult, expensive,
and generally unsuccessful. Research in Wisconsin, Illinois,
Minnesota, Missouri, and Iowa has shown clearly that fire
is not the enemy of the oak forest, but its friend.
Restoration of an Oak Savanna A principal
activity in oak savanna restoration is the "daylighting"
of oaks, removing trees that are crowding out the oaks. The
bur oak in the above photo is a good example. This oak, whose
age is estimated at about 200 years, was virtually invisible
when restoration work began, being completed crowded with
elm, cherry, and buckthorn. Once daylighted, the lower branches
could thrive again, and the tree has been saved for posterity.
Note also the other open-grown oaks farther up the hill. The
open-grown oak is a handsome specimen which has often survived
from the period before European settlement.
Oak Savanna Locations at Pleasant Valley Conservancy
The savanna habitats at Pleasant Valley are in the areas where
fire would have been most common. These include the south-facing
slope and the part of the ridge top that is nearest to the
south slope. Because the south-facing slope receives intense
sunlight, it is much drier (more xeric) than the north-facing
slope. Historically, the whole south slope was prairie, and
fire would have spread quickly up into the savannas.
Digital photos taken with a fish-eye lens have been used
to measure the openness of the canopy. The principal savanna
area is on the upper south slope and the ridge top, where
many large open-grown bur oaks are present. In this savanna
zone, the canopy ranges from about 25 to 70% cover. The oak
woods, where there are no open-grown oaks, has canopy cover
around 90%.
At Pleasant Valley Conservancy, we have oak savannas of two
types, bur oak and white oak. The bur oak savanna is predominantly
on the top of the ridge, on the dolomite cap. The white oak
savanna is mainly on the Jordan sandstone just below the dolomite.
As a result of restoration, the full magnificence of the oak
savanna can be appreciated. A hike along the upper ridge trail
takes one through the heart of our oak savanna.
Canopy cover analysis of Pleasant Valley Conservancy savannas. For the past two years, as one of our winter projects, we have been setting up a database of all the trees at Pleasant Valley Conservancy. Each tree 10 inches diameter or above has been given a permanent number marker, has been measured, its species determined, and its location recorded by GPS. Using these data, it is possible, using nomographs developed by the U.S. Forest Service, to calculate the percent canopy cover of any stand. The table below gives the data for all of the savanna areas, and for some of the oak woodland areas.
Needed for the calculations are the basal area (in square feet) per acre, and the number of trees per acre. These two measures are marked on the X and Y axes on the Forest Service graph, and the canopy cover in percent is read off the nomograph.
As the table shows, those sites that are open savanna have fairly low crown covers, whereas the oak woodland sites have high or complete crown cover. In between the open savannas and oak woodlands are a few sites that are best called "closed savanna".
When selecting species to plant in the understories of a site, it is important to consider the canopy cover of that site, since some plant species thrive in sunnier sites than others. Thus, understanding the canopy cover is useful in planning seed mixes for various sites. |
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Basal area calculations for Pleasant Valley Conservancy savanna areas
Based on tree database. Areas determined from ArcGIS.
Sorted by basal area per acre
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Management Unit |
Basal area per acre |
Trees per acre |
Crown cover, percent (see footnote for method) |
Interpretation |
2 (South slope) |
3.1 |
0.7 |
0 |
Prairie |
3 (South slope) |
3.5 |
0.6 |
0 |
Prairie |
22 (East Basin) |
15.6 |
14.6 |
13 |
Prairie |
18 (Bur oak savanna) |
21.3 |
8.7 |
19 |
Open savanna |
12A (White oak savanna) |
23.3 |
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~21 |
Open savanna |
Triangle (Remnant savanna) |
24.2 |
30.6 |
~25 |
Open savanna |
6 (South slope) |
26.8 |
15 |
24 |
Open savanna |
11D/11A (Remnant savanna) |
28.4 |
13.4 |
33 |
Open savanna |
14 (Transitional) |
32.4 |
32 |
~40 |
Open savanna |
10 (Bur oak savanna) |
32.7 |
16.2 |
31 |
Open savanna |
12B/11B |
35 |
21.1 |
36 |
Open savanna |
5 (Upper south slope) |
37.6 |
17.1 |
38 |
Open savanna |
7 (South slope: border between prairie and savanna) |
39.1 |
19.7 |
42 |
Open savanna |
20 (Border between savanna and woodland) |
39.7 |
37.8 |
48 |
Open savanna |
21 (Border between savanna and woodland) |
40.2 |
35.8 |
48 |
Open savanna |
8 (Bur oak savanna) |
46.8 |
23 |
52 |
Closed savanna |
19B (Edge of north woods) |
48.5 |
26 |
56 |
Closed savanna |
19CDE (Edge of north woods) |
50.3 |
32.1 |
59 |
Oak woodland |
13 (Edge of north woods) |
55.6 |
57.8 |
58 |
Oak woodland |
9 (Upper south slope) |
60 |
21.3 |
67 |
Oak woodland |
23 (Top of south slope) |
66.4 |
34.8 |
84 |
Oak woodland |
13A (Edge of north woods) |
82.3 |
80.4 |
100 |
Oak woodland |
17 (North woods) |
92.6 |
72.6 |
~100 |
Oak woodland |
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Footnote: Crown cover estimated from the nomograph published by Law, Johnson, and Houf 1994. Technical Brief No. 2. St. Paul, MN. U.S. Dept. of Agriculture, Forest Service, NorthCentralForest Experiment Station. TB-NC-2 Can be downloaded from http://www.treesearch.fs.fed.us/pubs/10999. The input data required were Basal area per acre and tree density per acre. Only trees greater than 4” diameter were measured. |
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